Sakly 2, Khémais Ben Rhouma 2, *Hichem Alimi 1, 2 , Zouhour Bouoni 3, Anwer Feriani 3, Najla Hfaeidh 3, Mohsen, 1 Research unit of Macromolecular Biochemistry and Genetic, Faculty of Sciences of Gafsa, University of Gafsa, 2112 Gafsa, Tunisia. 2 Laboratory of Integrated Physiology, Faculty of Science of Bizerte, University of Carthage, 7021 Jarzouna, Bizerte, Tunisia. 3 Laboratory of Animal Ecophysiology, Faculty of Science of Sfax, University of Sfax, 3018 Sfax, Tunisia., Tunisia